Had the same questions for a long time; here's the answer provided to me by a few trusty sources.
Grinding actions deflects to some extent, more and less depending on the local surface hardness and cooling action. This causes a slightly uneven surface, but avoidable in today's production environment.
The stationary surface called a way, and the moving surface called a slide: For box way systems, the way is usually ground (and hardened) while the slide is ground, scraped, flaked or frosted. Even when Turcite or Moglice is used for the slide, it is flaked. the purpose of scraping/flaking the slide is to prevent hydraulic locking of the slide and way. If they were both ground and left as-is, the slide would jerk and lurch as it moved plus the distance between the slide and way changes with slide velocity. For a mill saddle, it'll make irregular tooling marks and a sloped, stepped or poorly planed surface.
The next time I rebuild a machine tool, I'll be using Moglice or Turcite on the slide, rather the a full rescrape on both way and slide. I'll just scrape or grind the way.